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What do we know about the phonetic distinctions between the 甲類 and 乙類 syllables in 上代特殊仮名遣い?

上代特殊仮名遣い【じょうだいとくしゅかなづかい】 is a Nara-period practice in which two distinct versions of certain syllables (called 甲類【こうるい】 and 乙類【おつるい】, and denoted by subscript 1 and 2 in Latin script) were ...